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How Evolution Works Genes, Geography and Sex Early Idea: Lamarckian Evolution    Fossil record indicated increasing complexity Reasoned traits are passed via use and disuse e.g. the giraffe’s neck Step 1: Raw Material Genes are raw material  Forms of genes = alleles  Polygenic vs. Single gene trait  Selection on a Single Gene: The Peppered Moth  Before Industrial Revolution  After Industrial Revolution  Single gene controlling a trait will have only a few (usually 2 or 3 phenotypes) Variation and Selection Variation from two sources 1) New mutations = new allele types 2) Gene shuffling = new allele combinations  Any change in allele frequency = Evolution  Peppered Moth Simulation  Polygenic Traits  More than one gene controls a trait Selection and Changing the Norm Most traits are polygenic  The normal trait is the average or mean in the population  Selection changes the mean, usually lowers variation  Selection will adjust mean  Stabilizing Selection Disruptive Selection  Can lead to new species Does Evolution Ever Stop? YES, but only if the following conditions are met 1) Random mating 2) Large Population 3) No movement in or out of population 4) No Mutations 5) No Natural Selection  Hardy Weinberg Equilibrium  How New Species Are Formed  Two populations must become reproductively isolated Behavioral Isolation  Groups differ in mating, feeding, sleep/awake cycles so members of populations do not meet to mate Rana aurora (Redlegged frog)  Breeds in fast moving streams Rana catesbiana (Common bullfrog)  Breeds in still ponds Geographic Isolation Geological Time and Evolution  First fossils 3.5 bya Stromalites  550 mya = Cambrian Explosion (by fossil evidence) Dating Fossil Age  Relative Dating   Radioisotope Dating Known decay times of isotopes can be used Extinction and Adaptive Radiation  Most species cannot adapt  Those that can radiate into open niches Evolution Patterns Convergent Evolution Coevolution