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Chapter 29-33 Plant Unit Importance Evolution to land plants-classification Plant structure and function Plant Reproduction Plant Responses Plants and People Tree Man -Actually, this is believed to be caused by the Human Papilloma Virus (HPV) -Creepy though! Plants and People (really)  Plants are used as food  Plants are used for medicine  Plants are used for clothing and fabric dyes  Plants are used as fuels  Plants are also used for ornamental purposes, improving erosion, reducing noise, providing habitats for wild animals, acting as windbreaks, providing shade, and moderating temperatures  See page 566, Figure 29-3…good “Nonfood” list of plant uses Plants and the Environment  Plant Ecology – plant interaction in environment  Photosynthesis  Provide organisms with inorganic nutrients  Formation/maintenance of soil  Plant and animal interactions  Pollination  Size, shape, color attract pollinators  Plant and microbe interactions  Fungi – Mycorrhizae = symbiotic relationship betw. Plants and fungi  Bacteria – nitrogen fixing = take gas and fix so plants can use  Plant protection – prevents new plant pests  Harmful plants – can cause illness, rashes, allergies, and sometimes death Evolution of Land Plants  Changes in Nutrient Absorption  Vascular Tissue – transports water and dissolved substances  Xylem-water and inorganic nutrients in one direction  Phloem- organic compounds in any direction based on need  Reproduction by spores and seeds – keep reproductive cells from drying out; can send structures via air  Prevention of Water Loss – cuticle is waxy; prevents water loss but also keeps out carbon dioxide FYI: *Buoyancy of water provides physical support for algae Lignin is an evolutionary trait that hardens the cell walls of plants allowing them to have strength to stand upright Classification – Page 580, Figure 30-1  Nonvascular – no true vascular tissue nor true roots, stems, or leaves  Vascular, seedless – vascular tissue and true roots, stems, and leaves (ferns)  Vascular, seed – vascular tissue and true roots, stems, and leaves; also produce seeds for reproduction (4 phyla of gymnosperms, 1 phyla of angiosperms) Gymnosperms  Include pine trees  Produce seeds that are not enclosed in fruits Angiosperms  Flowering plants  Produce seeds within a protective fruit Alternating Life Cycles  2 phases  1st = diploid (2n); sporophyte plant; produces spores  2nd = haploid (1n); gametophyte plant; produces egg and sperm  Alternation of generations = alternates between the gametophyte phase and sporophyte phase Nonvascular Plants  “Bryophytes”  Do not form true roots, stems, and leaves  Usually grow on land near streams and rivers  Seedless; produce spores  Mostly terrestrial  Have alternation-of-generations life cycle  Very small  Need water to produce sexually (sperm must swim)  Phyla: Bryophyta, Hepatophyta, and Antherocerophyta Phylum Bryophyta  Pioneer plants = mosses; often first species to inhabit a barren area  Carpet moss  Moss gametophytes anchored by non-vascular rhizoids  Rhizoids anchor moss and absorb water and inorganic nutrients  Sphagnum (peat moss) – partially decomposed plant matter; some countries used as fuel; widely used to enhance the water-retaining ability of potting and gardening soils. Phyla Hepatophyta and Anthocerophyta  Hepatophyta – liverworts; grow in moist, shady areas  Lie close to ground  Most have thin, transparent leaflike structures arranged along a stemlike axis.  Anthocerophyta – hornworts; resemble liverworts; grow in moist, shady areas.  Similar characteristic to algae: each cell usually has a single large chloroplast rather than numerous small ones Changes in Nutrient Absorption  Land Plants  Algae  Absorb resources from surrounding water  Vascular Tissue transportation  Air  Light and carbon dioxide  Shoots/leaves  Soil  Water and mineral nutrients  Roots Vascular Plants  Vascular Seed Plants  Phylum Cycadophyta  Cycads  Seedless Vascular Plants  Phylum Psilotophyta  Whisk Ferns  Phylum Lycophyta  Club Mosses  Phylum Sphenophyta  Horsetails  Phylum Pterophyta  Ferns  Phylum Ginkgophyta  Ginkgoes  Phylum Coniferophyta  Conifers  Phylum Gnetophyta  Gnetophytes like Ephedra  Phylum Anthophyta  Largest phylum of plants  Over 240,000 of flowering plant Monocots vs Dicots Specialized Plant Cells – See pg 599  Parenchyma – loosely packed; cube-shaped  Involved in many metabolic functions (photosynthesis, storage of water and nutrients, and healing)  Bulk of nonwoody plants (ex. Flesh of apple)  Collenchyma – irregular shape; thick walls  Usually grouped in strands  Support regions of plant that are still lengthening (ex. Celery stalk)  Sclerenchyma – thick, even, rigid cell walls  Support and strengthen plant  Usually dies at maturity, providing support (ex. Gritty texture of pear fruit) Tissue Systems – Review Table 31-1  Dermal – outside covering of plants; surrounds ground tissue  Ground – consists of all 3 types of cells; surrounds vascular tissue system; storage, metabolism, and support  Vascular – transport and support; xylem and phloem; Growth in Meristems  Growth where cells continuously divide  Apical meristems – located at tips of stems and roots (remember the onion root tip?)  Some monocots also have intercalary meristems = above bases of leaves and stems; allow rapid regrowth from damage  Lateral meristems – gymnosperms and most dicots have this which allows roots and stems to increase in diameter  Vascular cambium – betw. xylem and phloem…produces additional vascular tissue  Cork cambium – located outside phloem…produces cork; cork cells replace epidermis in woody stems and roots; cork cells are dead cells that provide protection and prevent water loss  Primary Growth – in length; apical and intercalary  Secondary Growth - in diameter; lateral Roots  Taproot – when first root becomes the largest root; carrots  Fibrous root system – numerous small roots; many monocots…grass  Adventitious – specialized roots that grow from stems and leaves; prop roots of corn (helps keep stems upright) Root Structure  Root cap – covers apical meristem  Root hairs - extensions of epidermal cells (increase surface area = increase plants ability to absorb water)  Most form partnerships with mycorrhizal fungi = also increases surface area for absorption Just another FYI Cross section of monocot and dicot roots Stems  See page 609…  Shoot system and root system  Segmented stems = internodes  At each end of internode = node; contain 1+ leave and lateral bud (capable of developing into a new shoot) *Read over page 610 to review primary and secondary growth in stems Cross section of mature woody stem Bark – protective outer covering Heartwood – darker wood in center of tree Sapwood – lighter wood near outside Springwood/Earlywood – When water is plentiful; vascular cam. forms new xylem tissue (wide and thin-walled Summerwood/Latewood – When water is more limited; vasc. cam. forms small, thick cells. Annual Ring – abrupt change between small summerwood cells and following year’s large springwood cells. *Annual rings often do not occur in tropical trees because of their uniform year-round environment Stem functions  Source to sink –  CHO are “translocated” through the plant  Source = where CHO are made or stored  Sink = where the CHO are transported to be stored  EX. In most plants…CHO from leaves to roots …to the shoot apical meristems, and to the developing flowers or fruits PressureFlow Hypothesis Page 612 -Movement in phloem -CHO are actively transported into the sieve tubes -water is also transported in by osmosis -+ pressure builds up -At “sink”, process reverses Cohesiontension theory -Water is pulled up the Stem xylem b/c water molecules are attracted to one another (cohesion) How roots absorb water and minerals and then move to leaves  Trace the path of water and minerals from the roots to the leaves with the use of xylem  Trace the path using a simple concept map, isolating key points Types of Leaves Leaf Tissue Another Crosssection Tissue Systems  Dermal – epidermis  Single layer  Coated by cuticle  Water, oxygen, and carbon dioxide enter and exit through stomata  Mesophyll layers –  Leaf = ground tissue made of chloroplast-rich parenchyma cells  Palisade medophyll  Spongy mesophyll  Vascular Tissue Venation Moisture Maintenance  Must maintain a watery internal environment  Developed a cuticle – waxy coat that does NOT permit gas exchange  Stomata permit gas exchange during certain times of the day Plants acquire nutrients from soil and air  Three sources of a plant’s mass:  Air – carbon and oxygen = used in photosynthesis = generates sugar which are the building blocks of other organic molecules  Water – absorbed by plant – supplies the hydrogen for photosynthesis; solvent for transport of other molecules; makes up about 80-85% of a nonwoody plant’s mass  Soil – source of inorganic nutrients (minerals) Mineral requirements of plants  Most plants need 17 chemical elements to complete their life cycles:  3 are not obtained as minerals: C, O, and H  Mineral nutrients absorbed in ionic form from the soil  6 of mineral nutrients plants require in greatest abundance: Nitrogen, Sulfur, Phosphorus, Potassium, Calcium, Magnesium Path of N from atmosphere to plant’s roots  N is often in limited supply  Used to produce proteins, nucleic acids, and hormones  Makes up 80% of atmosphere, however atmospheric Nitrogen is a gaseous form of nitrogen that plants can’t use  Plants must absorb from the soil in the form of mineral ions Path of N from the atmosphere to plant’s roots  N must first be converted to ammonium ions or nitrate ions  Certain species of soil bacteria convert atmospheric nitrogen to ammonia (nitrogen fixation)  Ammonifying bacteria contribute ammonia to the soil by breaking down feces and dead leaves  Nitrifying bacteria convert ammonium ions to nitrate ions  Nitrogen-fixing bacteria live freely in the soil, however, some plants house their own (legumes)  Bacteria found in lumps on the legumes’ roots (root nodules)  Relationship benefits legumes and nitrogen-fixing bacteria  Bacteria fix nitrogen  Plant provides CHO and other organic nutrients to bacteria Fertilizer contents  Comercially produced; contain minerals  Usually enriched in nitrogen, phosphorus, and potassium (3 most deficient in farm and garden soils)  “10-12-8” three-number code in fertilizer; 10% Nitrogen, 12% phosphorus, and 8 % potassium  Not stored in soil for later use, though  Usually wasted; taken from soil by rainwater and irrigation = may pollute groundwater, streams, and lakes  Rotate crops reduces N fertilizer use  Mulch and manure release minerals more gradually Plant Reproduction  Life cycle of mosses  Life cycle of ferns  Life cycle of conifers Life cycle of mosses Production of one type of spore = homospory Life cycle of mosses = homosporous alternation of generations Male Reproductive structure Dominant generation = gametophyte Female Reproductive Structure Life cycle of ferns Dominant Generation = Sporophyte Similar to moss…most are homosporous Life cycle of conifers Microspores = male grow into male gametophytes Megaspores = female grow into female gametophytes Heterospory = production of different types of spores… Heterosporous alternation of generation Male cones of a pine release huge numbers of pollen grains Flower Parts Male reproductive structure = stamens…made up of an anther and filament Female reproductive structure = pistil…made up of a stigma, style, and ovary Flower pollination Fruit and seed dispersal  Dispersed by animals, wind, water forcible discharge, and gravity  Fruit = a matured ovary  Simple = formed from one pistil of a single flower  Aggregate = formed from several pistils of a single flower  Multiple = formed from several flowers growing together  Seed coat = protective coat surrounding embryo of seed  Cotyledon = seed leaf (part of embryo)…dicot = 2 cotyledons  Embryo  Radicle = embryonic root  Hypocotyl = stem between the cotyledons and radicle  Epicotyl = stem above the cotyledons Structure of seeds Conditions for seed germination Water Oxygen Temperature Vegetative propagation Use of vegetative structures to produce new plants Rhizomes, bulbs, and tubers Plant Horomones – see Page 648, Table 33-1  Auxins = involved in plant-cell elongation, apical dominance, and rooting  Gibberellins = stimulates elongation growth  Ethylene = responsible for ripening of fruit (hang green bananas above apples on counter…bananas ripen; ripen peaches in brown paper bag b/c they release ethylene and stimulate ripening)  Cytokinins = promote cell division in plants (remember “cytokinesis”)  Abscisic Acid = inhibits other hormones (causes closure of plant’s stomata in response to drought) Plant Movement Also referred to as “gravitropism” Chemotropism = a plant’s growth response to a chemical change (growth of pollen tube) Nastic Movements Photoperiodism  Plants response to changes in the length of days and nights  Critical Night Length = plant’s specific requirement for darkness; regulates flowering  SDP = short-day plants; flowers when the days are short and the nights are longer  LDP = long-day plants; flowers when the days are long and the nights are shorter  DNP = day-neutral plants; not affected by day length Phytochrome = bluish, light sensitive pigment that allows plants to monitor changes in day length Vernalization  Low-temperature stimulation of flowering  Important for fall-sewn grain crops  Farmers often use this to grow and harvest their crops before a summer drought sets in Fall colors Caused by photoperiodic response and by temperature response Longer nights = leaves stop producing chlorophyll…chlorophyll degrades and carotenoids become more visible (hidden by more abundant chlorophyll