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How many species of spiders do you see? Vocabulary so we are all on the same page…  Species: group of organisms that breed with one another and produce fertile offspring  Share a common gene pool, largest gene pool possible  Speciation – formation of a new species  The shared gene pool is eventually split Reproductive Isolation  Reproductive Isolation – when members of two populations can no longer interbreed and produce fertile offspring  New species form when organisms are reproductively isolated.  REPRODUCTIVE ISOLATION is the KEY to SPECIATION! First a little story about our friend the squirrel… Geographic Isolation  Populations become The Grand Canyon separated by geographic barriers such as rivers, mountains, or other bodies of water. Abert Squirrel Kaibab Squirrel The Gene Pools must separate to become a new species  There is no gene flow between the populations AND  Over time, the populations respond to new variations and natural selection as separate units Speciation  Reproductive Isolation  Geographical Isolation  Temporal Isolation  Behavioral Isolation The squirrels Reduction of gene flow Temporal Isolation  Temporal Isolation – occurs when two or more subspecies reproduce at different times Day Blooming/Night Blooming Eastern and Western Spotted Skunks Speciation  Reproductive Isolation  Geographical Isolation  Temporal Isolation  Behavioral Isolation The squirrels The Skunks Behavioral Isolation  Behavioral Isolation –capable of interbreeding BUT have different reproductive strategies Western Meadowlark Eastern Meadowlark Speciation  Reproductive Isolation  Geographical Isolation  Temporal Isolation  Behavioral Isolation The squirrels The Skunks The Meadowlarks What type of Isolation is under way?  http://www.pbs.org/wgbh/nova/beta/evolution/evolution-action-salamanders.html How many species of spiders do you see? THE HAPPY FACE SPIDERS! Look different but CAN interbreed! What about the ligers and mules? Are there any other gray areas for the definition of a species? Testing the Speciation Hypothesis in nature  1995 Hurricane brought the Iguana iguana species to a new island in the Caribbean  Will it go extinct?  Will the gene pool remain unchanged?  Will it respond to new selection pressures and become reproductively Isolated? Testing the Speciation Hypothesis Artificially  Separate cages act as what?  Develop food preferences  Mate and lay eggs on different food sources Can lead to the start of what? Test your understanding Explain how geographical and reproductive isolation work together with mutations to cause variation within a species and possibly speciation.