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					Speciation What came first, the adult alligator or the alligator embryo in an egg? Anagenesis  Speciation in Time   Directional selection causes species to change so much their ancestors would be considered different species (they would no longer recognize their ancestors and breed with them). No net increase in species number. Cladogenesis  Two Species Descend from One   Usually two populations get separated and evolve differences.  Sometimes one stays the same, or similar, and another changes especially if a few individuals end up in a different place. Occasionally speciation can happen in an intact population. Speciation Allopatric Cladogenesis  Speciation by Geographic Isolation  Two populations separated by a barrier evolve in separate directions until they can no longer interbreed  Two species result. Allopatric Speciation Tree frogs are a single population. Allopatric Speciation The formation of a river may divide the frogs into two populations. Allopatric Speciation Over time, the divided populations may become two species that may no longer interbreed, even if reunited. Sympatric Cladogenesis   No geographic isolation Not many animal examples   ring species Polyploidy in plants Sympatric Speciation  Ring species Ensatina in California The Tempo of Speciation   Gradualism - slow and steady change in species. Punctuated equilibrium - slow evolution punctuated by short events of rapid evolution. Polyploidy   A duplicate chromosome in humans result in conditions like Down’s Syndrome. In plants it doesn’t seem to matter much at all and can sometimes result in a new species with twice or more the number of chromosomes as before.  Haploid, Diploid, Triploid, Polyploid Polyploidy - The Spartina Story   Before 1830 Spartina maritima found in British salt marshes. S. alterniflora arrived in 1829 from boats from America. Polyploidy - The Spartina Story   Two species hybridised produced a sterile diploid plant S. townsendii. Two chromosomes from different parent species could not pair up in meiosis. Spread widely through asexual reproduction via underground stems. Polyploidy - The Spartina Story  1829 chromosome doubling occurred (complete non-disjunction) producing a tetraploid plant S. anglica.   With two sets from each of the original parents now possessed by the plant, meiosis and sexual reproduction became possible again. It is very fertile and vigorous, and has replaced nearly all of the other species across England. Polyploidy  Bread Wheat selectively bred by crossing two species. Allometry   Rate of growth of body parts. Can change due to single mutations. Allometry Allometry - Paedomorphosis  Sexually mature juveniles. Allometry - Paedomorphosis  Or juvenile appearing adults? Homeotic gene mutations  Hox gene mutations can change phenotype dramatically.