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Cranial nerves: Are 12 pairs of nerves connected directly to the brain (to the brain stem) except the 1st pair. Theses nerves characteristic by : 1-it not has two roots like spinal nerves. 2-Some are sensory ,some are motor and some are mixed. 3-All the cranial nerves enervated the head except the vagus extended to the thoracic and abdomen . Functional compound of the cranial nerves : 1-The somatic and visceral efferent impulses . a-The general visceral efferent fibers carry motor impulses to skeletal muscles . b-The general visceral efferent fibers ( autonomic ) supply glandular tissue and smooth muscles. 2-The somatic and visceral afferent impulses : a-the general visceral afferent fibers ( interceptive ) convey impulses from visceral , gland , blood vessels and mucous membrane. b-The general somatic afferent fibers ( incoming ) : transmits impulses which receive stimulation outside world and supply the body with information concerning its environment. 3-The special visceral efferent ( SVE ) fibers : which supply the muscles derived embryologically from and related to the branchiomeres and theses muscles are striated and are mainly associated with visceral activities of the digestive and respiratory system. 4-The special visceral afferent ( SVA ) fibers: are the fibers which innervated taste buds and olfactory mucosa . 4- The special somatic afferent ( SSA ) fibers : the vision , hearing and equilibrium are special sense relate to somatic activities . The cranial nerves are : I. 1ST cranial nerve : Olfactory nerves are sensory for smelling . II. 2nd cranial nerve : Optic nerves are sensory for sight . III. 3rd cranial nerve: Oculomotor nerves are motor for movement of eye muscles . IV. 4th cranial nerve: Trochlear nerves are motor for movement of obliqus dorsalis muscle of eye . V. 5th cranial nerve : Trigeminal nerves are mixed nerves . VI. 6th cranial nerve : Abducent nerves are motor nerve for the movement of rectus lateralis and retractor oculi muscles of eye . VII. 7th cranial nerve : facial nerves are mixed . VIII. 8th cranial nerve : Auditory nerve or vistibulocochlear nerve are sensory responsible for hearing and balance . IX. 9th cranial nerve glossopharyngeal nerve are mixed . X. 10th cranial nerve : Vagus nerves are mixed . XI. 11th cranial nerve :Accessory nerves are motor for movement of brachiocephalicus and trapizeus muscles. XII. 12th cranial nerve :Hypoglossal nerves are motor for the movement of lingual muscles. 1ST Cranial nerve ( Olfactory nerves): 1-the olfactory nerves of smell are represented by a number of bundles of nerve fibers in mucous membrane of dorsal concha and nasal septum . 2-it passes the foramina of cribriform of the ethmoid bone. 3-they end in the olfactory bulbs where they synapse . 4-the fibers not connect with brain stem . 5- They are not having ganglion . 2nd Cranial nerve : Optic nerves are sensory for sight . a- Its arise from ganglion layer of the retina (ganglion cell layer). The nerve fibers of retina converge towards the optic papillae or disc as special somatic afferent fibers and pierce the choroids and sclerotic coat to emerge as the optic nerve. b- The nerve course caudally toward the brain and passes through the optic canal and joins the opposite nerve to form the optic chiasma. c- From the optic chiasma the fibers are continued in the optic tract to lateral geniculate bodies and mid brain. d- The number of nerve fibers in the optic nerve is related to importance and activity of the visual system in particular species. 3rd Cranial nerve: Oculomotor nerves: aAre motor nerves to the ocular muscles and supply all the muscle of the eye except the obliqus dorsalis, rectus lateralis and retractor oculi. b- It also contains autonomic fibers for the ciliary muscles . c- It arise b from the brain stem medial to the cerebral crura. d- It leaves the cranial cavity by way of the orbital fissure. e- It divided into two branches : 1-The dorsal branch –supply the rectus dorsalis and levator palperbal superioris 2-The ventral branch—supply the ciliary ganglion and rectus medialis , ventralis and obliqus ventralis muscles of eye 4th Cranial nerve: Trochlear nerves A-are motor for movement of obliqus dorsalis muscle of eye . b- is the smallest of the cranial nerves , it arise from the rostral cerebral peduncle caudal to the caudal colliculus . c- It is only cranial nerve to emerge from the dorsal surface of the brain stem. d- It leaves the cranium cavity through a small foramen immediately above the orbital fissure or through the orbital fissure . 5th Cranial nerve : Trigeminal nerves are mixed nerves . a- It is connected with the lateral part of pons by large sensory root and smaller motor root. b- The sensory root connected with large trigeminal ganglion( semilunar ganglion ) which occupy the rostrolateral part of foramen lacerum , then from the rostral face of this ganglion arise three division ( ophthalmic , maxillary and mandibular nerves ) c- The motor root extends rostrally beneath the sensory root and the trigeminal ganglion , it is incorporated with the mandibular nerve. d- The trigeminal nerve divides into three branches( ophthalmic , maxillary and mandibular ( have two root sensory and motor ). A-Ophthalmic nerve: a- is the smallest of the three branches. b- Its purely sensory . c- It arise from the from the medial part of the rostral part of the trigeminal ganglion. D-it leaves the cranium cavity through the orbital fissure with third and 6 th cranial nerve and divides into three branches 1- Lacrimal nerve –supply the lacrimal gland and upper eyelid. 2-Zygomaticotemporal branch –it arise alone or with the lacrimal by common trunk, it supply the skin of the temporal region. In ruminant (in horned animals ) most of fibers of it course toward the horn as the corneal branches terminated in the lateral caudal surface of the base of horn and emerges from the zygomatic process of the frontal bone. 3-Frontal nerve : runs rostrally almost parallel with the obliqus dorsalis , within the orbital , then pass through the supraorbital foramen as the supraorbital nerve( with artery ), then ramifies in the skin of the forehead and upper eyelids . 4-Nasociliary nerve : pass rostrally and medially under the rectus dorsalis muscle and divided into two branches a-the ethmoid nerve : which is the continuation of the trunk inter the nasal cavity through the ethmoid foramen and ramifies in the mucous of the nasal septum and dorsal nasal concha . b-Infratrochlear nerve : ramifies in the skin nasal cavity ,it gives branches to the conjunctiva , lacrimal caruncle , Long Branch to the 3 rd eyelid and lacrimal duct and sac. B-Maxillary nerve : 1-it purely sensory and is much larger the ophthalmic . 2-it extends from the trigeminal ganglion in the middle cranial fossa in the large maxillary sulcus of the basisphenoid bone . 3- It emerges through the foramen rotundum and is continued in the infraorbital canal. It divides into follows : a-Zygomaticofacial branch—arise before the maxillary nerve reaches the pterygopalatine fossa, it pierce the periorbital and divided into two branches and ramify chiefly in the lower eyelid and the adjacent skin. B-Pterygopalatine nerve –is given off in the pterygopalatine fossa , it is broad and flat , and have several small ganglion. It divided into three branches. 1-Caudal nasal –enter the nasal cavity and divided into : ( medial branch to the mm of the nasal cavity , vomeronasal organ and hard palate; lateral branch to the mm and divided into the ventral nasal concha and to the middle and ventral nasal meatus. 2-Greater palatine nerve – is the largest branch, runs in the greater palatine canal and palatine groove , ramifies in the hard palate and gums. 3-lesser palatine nerve – is the smallest branch and ramifies in the soft palate. 4-Infraorbital nerve : is the continuation of the maxillary trunk , it passes the infraorbital canal and emerges through the infraorbital foramen and divided into (1- external nasal branch-supply nose , 2- internal nasal branch –supply upper and lower lips and nostrils, 3- maxillary labial branch –supply lip and cheeks and 4- along it course in the canal gives off maxillary alveolar branch which supply the teeth and alveolar peroisteum and gums ) B-Mandibular nerve : a- it formed by union of the two roots ( the large sensory) comes from the trigeminal ganglion and the motor root is the smaller part . b- It emerges from the cranium cavity through the oval notch of the foramen lacerum and passes between the wing of the basisphenoid bone and the muscular process of the petrous part of the temporal bone and when reaches to the lateral surface of the pterygoideus medialis muscle divided into two terminal branches (mandibular alveolar and lingual nerves). The mandibular nerve gives the following branches 1-the masseter nerve –ramifies the masseter muscle . 2-deep temporal nerve—two or three in number , supply the temporal muscle. 3-Buccal nerve-it continuous rostral in the submucous tissue of the cheek along the ventral border of the depress or labii mandibularis and divided into branches ramify I the mm and gland of the lip in the lip commissure. 4-auriculotemporal nerve : pass between the parotid gland and the neck of the ramus of the mandible , turn around it and divided into two branches (the transverse facial branch –ramify in the skin of cheek , the rostral auricular branch ) 5-mandibular alveolar nerve :arise with the lingual nerve by common trunk , then the two separated at the acute angle and the mandibular branch enter the mandibular foramen and course in the mandibular canal ,it emerges at the mental foramen as the mental nerve which terminating by divided into six to night mandibular labial and mental branch , which ramify in the lower lip and chin. Before enter the mandibular canal it gives the mylohyoid nerve , which supply the mylohyoid m and inside the canal gives the mandibular alveolar branches supply the teeth . 6- Lingual nerve : arise by common trunk with the mandibular alveolar nerve after separating directly toward the root of the tongue and give the sublingual nerve which supplies the mm of the tongue and the floor of the mouth. Note: The choradatympani: branch of facial joins the lingual nerve at the origin of the lingual. 6th cranial nerve: Abducent nerves a- Are motor nerve. b- arise from the brain stem behind the pons directly lateral to the pyramid , c- Cranially pierce the dura mater with 3rd cranial nerve and ophthalmic nerve and leaves the cranium cavity through the orbital fissure. d- In the orbit divided into two branches to rectus lateralis and retractor oculi muscles of eye. 7th cranial nerve: Facial nerves are mixed nerve. 1- Arise superficially from the lateral part of the trapezoid body behind the pons, 2-the facial nerve enter the internal acoustic meatus with the vesibulocochlear nerve at the base of the meatus separated from each other and facial nerve pass through the facial canal. 3-inside the canal , the nerve has an curvature called the knee and above the knee the geniculate ganglion are found and the choradatympani originate from it. 4-it gives many branches inside the canal , then reaches to stylomastoid foramen. 5-after emerge from the stylomastoid foramen pass between the guttural pouch and the parotid salivary gland , and receive the auriculotemporal nerve before it emerge outside . 6-it gives many branches outside the canal before ended by two terminal branches ( the dorsal buccal and ventral buccal nerves).