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Chapter 15 POPULATIONS POPULATIONS Population one species in one place at one time. Units of evolution – how scientists study and predict evolution. Demography statistical study of populations. Used to predict how the size of a population will change. What Drives Evolution: There are 5 Forces of Change Only natural selection makes a population better adapted (more fit) to its environment. Mutations Provide Raw Material For Evolution mutation at the gene level mutation at the chromosome level Mutations are usually neutral or harmful in their effects; only rarely are they beneficial. Mutations “Just Happen” Mutations occur at random without regard to whether they have a beneficial, neutral or harmful Mutations are effect. a randomly acting evolutionary forces. Gene Flow or Migration Gene flow – movement of genes from one area to another. Immigration & Emigration are key factors in the human gene flow. Gene flow in plants – wind-dispersed pollen moving between Monterey pines. Gene Flow Immigration: moving INto an area • My mother immigrated to America when she was 18 years old. Emigration : EXITing an area. • She emigrated from England after WWII. Gene Flow or Migration Genetic Drift Genetic Drift - random fluctuation in allele frequency between generations. The effects of genetic drift are pronounced in small populations. FYI A Genetic Bottleneck is a Form of Genetic Drift In a genetic bottleneck, allele frequency is altered due to a population crash. FYI Genetic Bottleneck – A Historical Case Note: A genetic bottleneck creates random genetic changes without regard to adaptation. A severe genetic bottleneck occurred in northern elephant seals. Other animals known to be affected by genetic bottlenecks include the cheetah and both ancient and modern human populations. Endangered Species Are in the Narrow Portion of a Genetic Bottleneck and Have Reduced Genetic Variation The Effect of Genetic Drift is Inversely Related to Population Size Large populations = small effects. Small populations = large effects. FYI Founder Effect A founder effect occurs when a small number of individuals from one population Another Variation of Geneticisolated Driftfrom the original one. found a new population that is reproductively FYI Founder Effect - Another Variation of Genetic Drift The South Atlantic island of Tristan da Cunha was colonized by 15 Britons in 1814, one of them carrying an allele for retinitis pigmentosum. Among their 240 descendents living on the island today, 4 are blind by the disease and 9 others are carriers. FYI - The Founder Effect Old Order Amish populations are derived from a few dozen colonists who escaped religious persecution in Germany in 1719 to settle in Pennsylvania. The community closed. frequencies in Amish are significantly different Allele and geneticisdisease from the German ancestral and the surrounding local populations. FYI - The Founder Effect Non-Random Mating Non-random mating; Mates are chosen – as in maintaining a dog pedigree Inbreeding is mating with close relatives. • increases the frequency of recessive genetic disorders. Natural Selection An adaptation (favorable trait) allows nature to choose who will have a better chance of surviving and reproducing. Natural Selection works on the phenotype not the genotype. It’s not natural – but this is one outcome of strong selection. Copy this Graphic Organizer Populations Can evolve through Gene Mutation Gene Flow Non-random Mating Gentic Drift Natural Selection End of Day One KEY FEATURES OF POPULATIONS 1. Size - how many individuals there are –effects the ability of the population to survive. A small population is more likely to become extinct: Why? Drastically affected by: -random events or natural disaster -inbreeding. Recessive traits are more likely to appear. reduced variability it’s harder to be adapted to changes. KEY FEATURES OF POPULATIONS 2. Density – how close together in a given area. –too far apart: may only rarely encounter one another resulting in little reproduction. too close together: resources may be scarce. KEY FEATURES OF POPULATIONS Population size is limited by: density-dependent factors density-independent factors Disease Competition Predators Parasites Food Crowding The greater the population, the greater effect these factors have. Ex. Black plague in the Middle Ages – more deaths in cities Volcanic eruptions Temperature Storms Floods Drought Chemical pesticides Major habitat disruption (as in the New Orleans flooding) Most are abiotic factors KEY FEATURES OF POPULATIONS 3. Dispersion –how are individuals arranged. Most common PREDICTING POPULATION GROWTH Nearly all populations will tend to grow exponentially as long as there are resources available. 4 basic factors affecting population growth: birth rate & Immigration (growth) (into) death rate & Emigration (decline) (exiting) r(rate of growth)=birth rate – death rate FYI Exponential Growth - Problem Suppose you work for the month of July (31 days) If you earned a penny on July 1stand your pay doubled each day; what would be your final paycheck on July 31st? Write down an estimate in your notes. Exponential Growth Day 1 - 1¢ Day 2 - 2¢ Day 3 - 4¢ Day 4 - 8¢ Day 10 - $5.12 Day 20 - $5,242.88 Day 31 - $10, 737,418.24 PREDICTING POPULATION GROWTH – The “J” Curve Exponential growth curve: As a population gets larger, it also grows at a faster rate. Maximum population growth under ideal circumstances. FACT: No population exhibits this type of growth for long. “S” Curve – Logistic Growth Population numbers stay around the carrying capacity. Carrying capacity (K): number of organisms that an environment can support. PREDICTING POPULATION GROWTH Two modes of population growth. The Exponential curve (J-curve) occurs The Logistic curve (S-curve) shows the effect when there is no limit to population size. of a limiting factor (in this case the carrying capacity of the environment). POPULATION GROWTH STRATEGIES 2 ways (Strategies) a population can grow: r strategists: “J” curve. insects, bacteria Exponential growth; then crash K strategists: “S” curve. mammals, fish, birds Live within the carrying capacity Who are “r” and “K” strategists? r strategists (J curve) Small organisms/some plants Rapid maturations Quick to reach reproductive age Require no parent nurturing Have periods of time where eggs/endospores can remain dormant K strategists (S curve) Larger mammals, fish, birds Require parenting Slower to grow & mature Few offspring Natural Selection Shapes Populations Natural selection acts on phenotype, NOT genotype. THEREFORE: selection against unfavorable recessive alleles is SLOW. Hardy-Weinberg Principle In 1908 G.H. Hardy and Wilhelm Weinberg showed that dominant alleles do not replace recessive ones. The Hardy-Weinberg Principle states: populations do not change unless evolutionary forces act upon them. Hardy-Weinberg Principle, con’t • An important way of discovering why real populations change with time is to construct a model of a population that does not change. • This is just what Hardy and Weinberg did. • Their principle describes a hypothetical situation in which there is no change in the gene pool hence no evolution. Hardy-Weinberg Principle • Hardy-Weinberg Principle: – Predicts allele frequencies in a population – True only for a stable population – Not applicable in “real” life. Some force of change is always at work. Hardy-Weinberg Principle • p² + 2pq + q² = 1 • Frequencies of individuals that are: • p² = homozygous dominant allele • 2pq = heterozygous • q² = homozygous recessive allele Stabilizing Selection Favors the Average Directional Selection Favors one extreme Disruptive Selection Favors both Extremes Natural Selection Shapes Populations NATURAL SELECTION DIRECTIONAL SELECTION STABILIZING SELECTION DISRUPTIVE SELECTION Favors one extreme phenotype Favors the average phenotype Favors both extreme phenotypes Possible reason: Predators can identify easier and eat the average type organism Crash Course populations http://www.youtube.com/watch?v=RBOsqmB QBQk&list=PL8dPuuaLjXtNdTKZkV_GiIYXpV 9w4WxbX&index=2&edufilter=uD4dsTAhNKX _EYoUsFAfPQ