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Ecological Concepts-2 selfguru.org 1 Outline Ecotone Niche Biome Biosphere Ecosystem Productivity Ecosystem Stability Modification of Ecosystem Evolution of Species Neo-Darwinism Succession- Ecological Succession Primary and Secondary Succession Autogenic and Allogenic Succession Autotrophic and Heterotrophic Succession selfguru.org 2 Outline Sere Trophic Level Energy Flow in Ecosystem Food Chain Food Web Ecological Pyramids- Pyramid of numbers, biomass and energy Pollutants and Trophic Level Bioaccumulation and Biomagnification Biotic Interaction Types of Biotic Interaction Eutrophication, Harmful Algal Blooms & Coral Bleaching Ecological efficiency Questions selfguru.org 3 Ecotone/ Eco clines Transition zone or junction between two diverse ecosystems along with biotic communities and abiotic condition change. It may be very narrow or quite wide. Zone of tension, having conditions intermediate to adjacent ecosystems. Progressive increase in species composition of incoming community and decrease in species of outgoing community. Well developed ecotones contain some organisms entirely different from adjoining communities. selfguru.org 4 Ecotone/ Eco clines Edge effect- Sometimes number of species and population density of some of species is much greater in ecotone than adjoining areas. Organism which occur primarily or most abundantly in this zone are known as edge species. i.e. Density of birds is greater in mixed habitat of ecotone between forest and desert. selfguru.org 5 Niche Unique functional role and position of species in its ecosystem including • where it lives, • what resources it uses, • how and when it uses resources, • how it interacts with other species. No two species have exact identical niches. Greater diversity of niche => More ecosystem stability as large number of pathways for flow of energy and less fluctuation of species population. selfguru.org 6 Niche Forest can support large number of plant species as they occupy different niches. Like tall trees, short trees, shrubs, bushes, grasses- because of their varying heights their requirements for sunlight and nutrients differ and so they can survive together (reduced competition) Most important resources in niches of plants is moisture, nutrient- phosphorus and nitrogen, While for animals, it is food and shelter. There are different types of niches like Habitat niche, Food niche, Reproductive niche, Physical and chemical niche. selfguru.org 7 Niche selfguru.org 8 Biome Terrestrial part of biosphere is divisible into enormous region called biome, which are characterised by climate, vegetation, animal type and soil type. Most important climatic factors are temperature and precipitation. (Heat and Moisture). selfguru.org 9 Biomes selfguru.org 10 Biosphere Biosphere is part of earth where life can exist. It represents highly integrated and interacting zone comprising of atmosphere, hydrosphere and lithosphere. Biosphere is absent at extremes of North and South poles, highest mountains and deepest oceans, since existing hostile conditions. Energy required for life within biosphere comes from Sun and nutrients come from air, water and soil. selfguru.org 11 Ecosystem Productivity Growth of energy or organic matter per unit time by autotrophs at trophic level through photosynthesis Depending factors Availability of solar radiation at trophic level; Efficiency of plants to convert solar energy into chemical energy; Availability of moisture Net Primary Productivity= Gross Primary Productivity (total) – Loss through respiration NPP= usable amount of energy at trophic level selfguru.org 12 Ecosystem Productivity NPP decreases as one moves from equator to pole NPP decreases with increase in size of animals NPP of terrestrial ecosystem is generally greater than NPP of aquatic ecosystem Order of NPP of terrestrial ecosystem– 1. Swamp and Marsh-Tropical Forest 2. Temperate Forest 3. Tropical Savannah 4. Agricultural Land 5. Temperate Grassland 6. Extreme Desert-Rocks-Ice Order of NPP of aquatic ecosystem1. Attached algae & Estuaries- Coral Reef 2. Continental Shelf 3. Open Sea selfguru.org 13 Ecosystem Stability Ecosystem stability is the ability of ecosystem to maintain its structure and function over long periods of time and despite disturbances. Ecosystem structure includes physical and geological structures of landscape, number and diversity of species present, population of size of those species, ways in which population interact. 1. 2. 3. 4. Ecosystem function refers to processes such as water and nutrient cycling and biomass productivity that ecosystem provides. selfguru.org 14 Ecosystem Stability Increase in one species is compensated by decrease in other species thus productivity is maintained. “Homeostasis”- Self regulating, natural ecosystem adjust changes to itself. Increase in stability => with increase in diversity and complexity of ecosystem Increase in stability => Decrease in competition as different resources can be used or use of same resources but at different time. selfguru.org 15 Modification of Ecosystem Destruction of natural/ original/ native species and introducing alien species which come in direct competition or bring diseases. Climate Change Chemical Fertilizers Atmospheric Pollution Depletion of Species Urbanization, Industrialization, Deforestation, Irrigation Overexploitation of resources selfguru.org 16 Evolution of Species Speciation Natural Selection Progressive evolution of Species 1. Genotype 2. Phenotype Neo-Darwinism Origin, growth and development of species through time Mutation Changes Reproductive Isolation Ecological Isolation selfguru.org Geographical Isolation 17 Evolution of Species Mutation- Spontaneous and abrupt evolution cause new gene to arise in population Reproductive Isolation- no exchange of specific gene from parent to off spring; Sexually receptive at different times; not attracted towards each other; Pollination mechanism fails; Not matching reproductive organs. Ecological Isolation- When ecologically-based divergent selection creates reproductive barriers between populations Geological Isolation- population of animals, plants, or other organisms that are separated from exchanging genetic material with other organisms of the same species. Typically geographic isolation is the result of an accident or coincidence selfguru.org 18 Neo-Darwinism Organisms produce more offspring than it can be supported by environment Members of same species shows variation because of either mutation (cause new gene to rise) or recombination (sexually reproducing population; meiosis and fertilization produce new combination of genes every generation) Only those genes that help organisms to adopt its environment are selected through natural selection. Those offspring which are suited to their immediate environment have more chance of surviving, reaching reproductive age and passing on suitable adaption. Evolution thus results in adaption and diversity of species. selfguru.org 19 Neo-Darwinism selfguru.org 20 Succession When ecosystem undergoes succession there is change in structure and productivity of ecosystem. Structure of ecosystem refers to complexity and diversity of ecosystem. Plant succession which is limited by water availability known as Xerarch succession on terrestrial ecosystem with low moisture. While, succession in aquatic ecosystem with high moisture known as Hydrarch succession. Succession will increase soil structure, composition and maturity which intend to increase relative stability of ecosystem. selfguru.org 21 Ecological Succession Entire process of directional and sequential change of biotic and abiotic component of ecosystem over period of time. Succession occurs when series of communities replace one another due to large scale destruction either natural or manmade. Succession is progressive series of changes which leads to establishment of relatively stable climax community. Succession is characterised by increased productivity, shift of nutrients from reservoirs, increased diversity, increased niche development and gradual increase in complexity of food webs. selfguru.org 22 Ecological Succession selfguru.org 23 Primary Succession Primary succession occurs on new site like bare or unoccupied areas, rocks, newly formed deltas, sand dunes, emerging volcano islands, glacial moraines (muddy area exposed by retreating glacier) Pioneer species- like microbes, lichens and mosses which invade first and colonise area. 1. These species are having high growth rate but short life span. 2. Pioneers take long time as soil has to be formed and over few generations of pioneers, they alter habitat condition and made it conducive to establishment of additional organisms. Serial species- Transitional species which succeeds pioneers and further helps in development of habitat condition. selfguru.org 24 Primary Succession Community of organisms continues to develop, it becomes more diverse and competition increases, but at same time new niche opportunities develops. Pioneer species disappear as habitat conditions change and invasion of new species progresses, leading to replacement of preceding community. Climax Species- Final stage of succession, more stable, mature and complex species which are having long life span and slow growth rate. selfguru.org 25 Secondary Succession Occurs when plants recognise an area in which climax community has been disturbed. Complete or partial destruction of existing community- mature or intermediate community may be destroyed by natural events like floods, hurricane, forest fire or by human interventions like harvesting, deforestation, overgrazing etc. Difference between primary and secondary succession is, secondary succession starts on well developed soil already formed at site. Thus, secondary succession is relatively faster as compared to primary succession. selfguru.org 26 selfguru.org 27 Autogenic and Allogenic Succession Autogenic succession- Succession is brought about by living inhabitants of that community itself. Allogenic succession- Succession is brought about by outside forces. selfguru.org 28 Autotrophic and Heterotrophic Succession Autotrophic succession- Succession in which initially green plants (autotrophs) are much greater in quantity. Heterotrophic succession- Succession in which initially heterotrophs are greater in quantity. selfguru.org 29 Sere Transitional stages of sequential change from one vegetation community to another. The whole sequence of communities that replace one another in a given area Maximum growth and development. Most adjustable and adaptable. selfguru.org 30 Trophic Level Trophic level of an organism is the position it occupies in food chain. It is point where food energy is transferred from one group of organisms to other. Trophic level interaction deals with how members of ecosystem are connected based on nutritional needs. selfguru.org 31 Energy Flow in ecosystem Flow of energy from producers to top consumers is called energy flow. Energy flow is unidirectional- always flows from lower to higher trophic level. Loss of some energy in form of unusable heat at each trophic level so that energy level decreases from first trophic level upwards. selfguru.org 32 Food Chain The chain of transfer of food energy from one group of organisms to other group in ecosystem. 1. Grazing food chain- begins with plants at base 2. Detritus food chain- starts from dead organic matter selfguru.org 33 Food Web Food web is when food chain is complicated by several overlapping and interconnected layers. Food web illustrates, all possible transfers of energy and nutrients among organisms in ecosystem. Richer the biodiversity, the longer and more complicated food chains and webs. selfguru.org 34 Ecological Pyramids- Pyramid of numbers Number of individuals in each trophic level is considered. Can be upright (Grassland ecosystem) or inverted (Tree ecosystem) selfguru.org 35 Ecological Pyramids- Pyramid of Biomass Total dry weight of all organisms at each trophic level at particular time is considered Biomass is measured in g/m2. Can be upright or inverted selfguru.org 36 Ecological Pyramids- Pyramid of Energy Energy pyramid reflects law of thermodynamics, with conversion of solar energy into chemical energy and heat energy being depicted at each transfer to another trophic level. Most suitable pyramid to compare functional roles of trophic level. Energy pyramid is always upward with large energy base at bottom. Energy pyramid helps to explain phenomenon of biological magnification- tendency of toxic substances to increase in concentration progressively at higher levels of food chain. selfguru.org 37 Ecological Pyramids- Pyramid of Energy selfguru.org 38 Ecological efficiency • The % of energy transferred from one trophic level to next varies between 5-20 % and is called ecological efficiency • An average figure of 10% is used known as ten percent law-total energy content of a trophic level in an ecosystem is only about 1/10th that of preceding level • Lost energy is used to catch , eat, digest, respiration, feces etc. selfguru.org 39 Pollutant and Trophic Level Non-degradable pollutants (POPs) like chlorinated hydrocarbons which can not be metabolised by living organisms, move through various trophic level in an ecosystem. Movement of these pollutants involves two processes: 1. Bioaccumulation 2. Biomagnification selfguru.org 40 selfguru.org 41 Bioaccumulation and Biomagnification Bioaccumulation- refers to how pollutants enter food chain; there is an increase in concentration of pollutants from environment to first organism in food chain. Biomagnification- refers to tendency of pollutants to concentrate as they move from one trophic level to next level. In order for biomagnification to occur, pollutants must be long-lived, mobile, soluble in fats, biologically active. selfguru.org 42 selfguru.org 43 Biotic Interaction Amensalism- (antibiotics) One species harm other species without itself getting affected. i.e. Bread mold fungi Pencillium produce penicillin, an antibiotic that inhabit growth of variety of bacteria, thus penicillium benefits from having greater availability of food as bacteria is removed. Predation- Predator capture, kill and eat the prey. i.e. Leopard, Tiger, Cheetahs as predator having speed, teeth and claws to hunt. Parasitism- One species is harmed while other is benefits. i.e. Parasites like bacteria, fungi, virus, roundworm live on host for food, nourishment and shelter while host is harmed. Neutralism- No net benefit or harm to either species. They do interact but do not affect each other. Relationship between two species inhabiting same space and using same resources but have no effect on each other. selfguru.org 44 Biotic Interaction Competition- Both species are harmed to some extend. Within species or interspecies competition for food, shelter, water, mate, space, resting site. Commensalism- One species benefits, while other is unaffected. i.e. Fish Remora attached to Shark. Remora is benefited as it gets protection, free ride and meal (shark left over food), while Shark is unaffected. Epiphytes- ferns, mosses, orchids. Mutualism- Both species benefit. Symbiotic- intimate relationship. i.e. Termites and Protozoa, Termites eat food but can’t digest while Protozoa digests and convert into sugar and uses sugar. Pollination mutualism, where pollinator gets food and plant has its pollen transferred for reproduction. selfguru.org 45 selfguru.org 46 Eutrophication Syndrome of ecosystem, response to addition of artificial or natural substances such as nitrates and phosphates through fertilizers, sewage etc. that fertilize aquatic ecosystem. Growth of green algae- physical identification of Eutrophication Some algae and blue-green bacteria thrive on excess ions and population explosion covers almost entire surface layer is known as algal bloom. This growth is unsustainable. Algal bloom covers surface layer and restricts penetration of sunlight; Oxygen level is reduced due to population explosion; New anaerobic conditions can promote growth of bacteria Clostridium botulinum which produces toxins– All these conditions leads to change in animal and plant population and degradation of aquatic ecosystem and habitat quality. selfguru.org 47 Eutrophication Algal bloom covers surface layer and restricts penetration of sunlight; Oxygen level is reduced due to population explosion; New anaerobic conditions can promote growth of bacteria Clostridium botulinum which produces toxins– All these conditions leads to change in animal and plant population and degradation of aquatic ecosystem and habitat quality. Effects of Eutrophication Change in Ecosystem Decreased Biodiversity New species invasion Toxicity selfguru.org 48 Harmful Algal Bloom When environmental conditions are favourable to development of algae and phytoplanktons, they can be multiplied rapidly and results in algal bloom. Bloom often results in colour change in water. Algal bloom can be any colour, but common ones are red or brown. These blooms are commonly referred as “Red Tides”. Most algal blooms are not harmful but some produces toxins and do affect aquatic organisms and humans. Toxins may affect surrounding air difficult to breathe. These known as Harmful Algal Blooms (HABs). selfguru.org 49 BOD,COD,DO 1. Biochemical oxygen demand (BOD, also called biological oxygen demand) is the amount of dissolved oxygen needed (i.e., demanded) by aerobic biological organisms to break down organic material present in a given water sample at certain temperature over a specific time period 2. Chemical oxygen demand (COD) is measure the amount of organic compounds in water. • Most applications of COD determine the amount of organic pollutants found in surface water (e.g. lakes and rivers) or wastewater, making COD a useful measure of water quality. selfguru.org 50 BOD,COD,DO 3. Dissolved oxygen measures the amount of gaseous oxygen (O2) dissolved in an aqueous solution. • Oxygen gets into water by diffusion from the surrounding air, by aeration (rapid movement), and as a waste product of photosynthesis. • Total dissolved gas concentrations in water should not exceed 110 percent. Concentrations above this level can be harmful to aquatic life. Fish in waters containing excessive dissolved gases may suffer from "gas bubble disease"; however, this is a very rare occurrence. The bubbles or emboli block the flow of blood through blood vessels causing death. External bubbles (emphysema) can also occur and be seen on fins, on skin and on other tissue. Aquatic invertebrates are also affected by gas bubble disease but at levels higher than those lethal to fish. • Adequate dissolved oxygen is necessary for good water quality. Oxygen is a necessary element to all forms of life. • Natural stream purification processes require adequate oxygen levels in order to provide for aerobic life forms. As dissolved oxygen levels in water drop below 5.0 mg/l, aquatic life is put under stress. • The lower the concentration, the greater the stress. Oxygen levels that remain below 1-2 mg/l for a few hours can result in large fish kills. selfguru.org 51 selfguru.org 52 Coral Bleaching Bleaching of coral occurs when densities of zooxanthellae decline and/or concentration of photosynthetic pigments within zooxanthellae fall. Major ecological causes of coral bleaching are Temperature (Rise or Fall) Solar Irradiance Subaerial Exposure Sedimentation Fresh Water Dilution Inorganic Nutrients Xenobiotics- concentration of various chemical contaminants such as Cu, herbicides and oil. Epizootics- Pathogens have been identified to cause translucent white tissues, a protozoan. selfguru.org 53 Adaptation • Each organism is adapted to its particular environment. • An adaptation is thus, “the appearance or behaviour or structure or mode of life of an organism that allows it to survive in a particular environment”. • Examples of basic adaptations that help animals and plants to survive in their respective environments. • Shape of bird’s beak. • The thickness or thinness of fur. • Presence of feathers and wings in birds. • Evergreen and deciduous nature of trees. • Presence and absence of thorns on leaves and stems. selfguru.org 54 Adaptation selfguru.org 55 Questions selfguru.org 56